Manchester United's big summer spending means that they now have the third most expensive squad in Europe, behind only neighbours Manchester City and Real Madrid.
United have spent £389.1m on Louis van Gaal's current squad, with City spending £408.1m and Madrid shelling out £428.5m, according to the CIES Football Observatory.
Van Gaal's squad cost over £100m more than Champions League winners Barcelona's, bought for £287m, and more than 50% more than Bayern Munich's, who arrived for £246m.
Morgan Schneiderlin and Memphis Depay both arrived at Old Trafford for respective £24m and £25m fees, while Anthony Martial arrived on deadline day for £36m.
Juan Mata cost £37m, Anders Herrera £28m, and Luke Shaw, Wayne Rooney and Marouane Fellaini all cost £27m each.